Introduction 1
Previous work 2
Geological setting 3
Tectonic history of Ganderia and Avalonia 4
The Iapetus Ocean 5
Ireland 6
The Leinster Terrane 7
Docking history 7
Th e Tramore area 7
Material 9
Stratigraphy and fauna 9
Age of the Tramore Limestone Formation 9
Tramore and Dunabrattin/Bunmahon sections 10
Tramore Shale Formation Tramore area 12
Tramore Limestone Formation type section at Barrel Strand, Tramore area 12
Lithology 12
Fauna associations 14
Tramore Limestone Formation other sections 14
Pickardstown sections (Ps) 14
Quillia Section 1 (QS1) 14
Quillia Section 2 (QS2) 14
Bunmahon Formation Bunmahon/Dunabrattin area 15
Dunabrattin Shale Formation Bunmahon/Dunabrattin area 15
Dunabrattin Limestone Formation Bunmahon/ Dunabrattin area 15
Lithology and fauna 16
Correlation 16
Depositional setting and ecology 17
Local events 17
Tramore Limestone Formation 17
Dunabrattin Limestone Formation 19
Palaeobiogeographical reconstructions 20
New reconstructions for the Iapetus Ocean 22
Ganderian terrane positions 22
Longitudinal separation 22
Latitudinal drift of the Ganderian-Avalonian segment 22
Brachiopod provinces 24
Faunal distribution 24
Darriwilian provinces and faunas 26
Th e Low-latitude Province on the Laurentian Platform 26
An early Scoto-Appalachian fauna 27
Th e Toquima-Table Head province 28
Th e mixed faunas of the Leinster Terrane and East Avalonia 31
Th e Leinster Terrane 31
East Avalonia 32
Th e Celtic province 32
Th e Baltic Province 34
Th e High-latitude Province 35
Darriwilian faunal migration patterns: implications on oceanic current confi guration 35
Migrations from the Laurentian margins and intraoceanic islands to the south IapetusTornquist region 38
Migrations within the south IapetusTornquist region 38
Th e Rheic gyre and the High-latitude Province 40
Th e Precordillera 40
Early Sandbian brachiopod provinces 41
Th e Low-latitude Province on the Laurentian Platform 41
Th e Scoto-Appalachian Province 42
Th e Anglo-WelshBaltic Province 43
Th e Leinster Terrane 44
East Avalonia and Anglesey 46
Baltica 47
The High-latitude Province 48
The Precordillera 48
Conclusions 48
Systematic palaeontology 50
Abbreviations 50
Repository 50
Statistical methods 50
Allometric growth 50
Phylum Brachiopoda Duméril, 1806 51
Subphylum Rhynchonelliformea Williams, Carlson& Brunton, 1996 51
Class Strophomenata Öpik, 1934 51
Order Strophomenida Öpik, 1934 51
Superfamily Strophomenoidea King, 1846 51
Family Strophomenidae King, 1846 51
Subfamily Strophomeninae King, 1846 51
Genus Tetraphalerella Wang, 1949 51
Tetraphalerella? sp. 52
Subfamily Furcitellinae Williams, 1965 53
Genus Dactylogonia Ulrich& Cooper, 1942 53
Dactylogonia costellata n. sp 53
Family Rafi nesquinidae Schuchert, 1893 55
Subfamily Rafi nesquininae Schuchert, 1893 55
Genus Colaptomena Cooper, 1956 55
Colaptomena auduni n. sp. 56
Colaptomena pseudopecten? (MCoy, 1846) 57
Superfamily Plectambonitoidea Jones, 1928 59
Family Plectambonitidae Jones, 1928 59
Subfamily Taphrodontinae Cooper, 1956 59
Genus Isophragma Cooper, 1956 59
Isophragma parallelum n. sp. 59
Family Leptellinidae Ulrich& Cooper, 1936 61
Subfamily Leptellininae Ulrich& Cooper, 1936 61
Genus Leptellina Ulrich& Cooper, 1936 61
Leptellina llandeiloensis (Davidson, 1883) 61
Subfamily Palaeostrophomeninae Cocks& Rong, 1989 62
Genus Glyptambonites Cooper, 1956 62
Glyptambonites sp. 62
Family Grorudiidae Cocks& Rong, 1989 63
Genus Grorudia Spjeldnæs, 1957 63
Grorudia grorudi Spjeldnæs, 1957 63
Family Leptestiidae Öpik, 1933a, emend. Cocks& Rong, 1989 64
Genus Leptestiina Öpik, 1933a 64
Leptestiina derfelensis (Jones, 1928) 65
Family Sowerbyellidae Öpik, 1930 66
Subfamily Sowerbyellinae Öpik, 1930 66
Genus Sowerbyella Jones, 1928 66
Subgenus Sowerbyella (Sowerbyella) Jones, 1928 66
Sowerbyella (Sowerbyella) antiqua Jones, 1928 66
Order Billingsellida Schuchert, 1893 70
Suborder Clitambonitidina Öpik, 1934 70
Superfamily Clitambonitoidea Winchell& Schuchert, 1893 70
Family Clitambonitidae Winchell& Schuchert, 1893 70
Genus Atelelasma Cooper, 1956 70
Atelelasma longisulcum n. sp. 70
Class Rhynchonellata Williams, Carlson& Brunton, 1996 73
Order Orthida Schuchert& Cooper, 1932 73
Suborder Orthidina Schuchert& Cooper, 1932 73
Superfamily Orthoidea Woodward, 1852 73
Family Orthidae Woodward, 1852 73
Genus Sulevorthis Jaanusson& Bassettt, 1993 73
Sulevorthis aff S. blountensis (Cooper, 1956) 73
Family Glyptorthidae Schuchert and Cooper, 1931 74
Genus Glyptorthis Foerste, 1914 74
Glyptorthis crispa (MCoy, 1846) 74
Family Hesperorthidae Schuchert& Cooper, 1931 77
Genus Hesperorthis Schuchert& Cooper, 1931 77
Hesperorthis leinsterensis n. sp. 77
Family Plaesiomyidae Schuchert, 1913 86
Subfamily Plaesiomyinae Schuchert, 1913 86
Genus Valcourea Raymond, 1911 86
Valcourea confinis (Salter, 1849) 86
Family Productorthidae Schuchert& Cooper, 1931 87
Subfamily Productorthinae Schuchert& Cooper, 1931 87
Genus Productorthis Koz?owski, 1927 87
Productorthis sp 87
Superfamily Plectorthoidea Schuchert& LeVene, 1929 89
Family Platystrophiidae Schuchert& LeVene, 1929, emend.
Zuykov& Harper, 2007 89
Genus Platystrophia King, 1850, emend. Zuykov& Harper, 2007 89
Platystrophia tramorensis n. sp. 89
Platystrophia aff P. sublimis Öpik, 1930 92
Suborder Dalmanellidina Moore, 1952 94
Superfamily Dalmanelloidea Schuchert, 1913 94
Family Dalmanellidae Schuchert, 1913 94
Subfamily Dalmanellinae Schuchert, 1913 94
Genus Howellites Bancroft , 1945 94
Howellites hibernicus n. sp. 94
Family Paurorthidae Öpik, 1933b 106
Genus Paurorthis Schuchert& Cooper, 1931 106
Paurorthis aff P. parva (Pander, 1830) 106
Superfamily Enteletoidea Waagen, 1884 110
Family Linoporellidae Schuchert& Cooper, 1931 110
Genus Salopia Williams in Whittington& Williams, 1955 110
Salopia gracilis Williams in Whittington& Williams, 1955 110
Order Pentamerida Schuchert& Cooper, 1931 111
Suborder Syntrophiidina Ulrich& Cooper, 1936 111
Superfamily Syntrophioidea Ulrich& Cooper, 1936 111
Family Porambonitidae Ulrich& Cooper, 1936 111
Genus Hibernobonites n. gen. 111
Hibernobonites filosus (MCoy, 1846) 112
Acknowledgements 115
References 115
Plates 110 122
Appendix A 142
Appendix B 148
